I have a theory about "404": He is Mr. E.
Their colour-scheme is the same (with the addition of blue in the case of "404"), "404" appears to have silver metal legs, and, aside from the three SoG Generals, no other Ninjago characters have numbers on their outfits (including the leaked Minifigs), so this is an odd stylistic choice for someone unrelated. Additionally, the 404 reference fits with the existing play on the word "mystery" - I.E. "mystery character not found".
Story-wise, Mr. E is the only Nindroid whose creator is unknown (Harumi's comment that he was "built to be a cut above the rest" precludes the producers' original plan for him to be Echo Zane), so he could be connected to Unagami/Prime Empire. Furthermore, even if Mr. E were not originally associated with Unagami, he could easily have created a backup of himself at Borg Tower before he was 'killed' (which would not be the first time that this has occurred - See Zane), and somehow ended up in the game.